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(July 12, 2014 at 8:17 pm)Lek Wrote: Evidence for an evolutionary process is obvious and plentiful, but of course, creating amino acids in a lab is far from creating a living creature. It also doesn't in any way rule out a creator. It may be true, but it's a lousy argument to use to try to deny the existence of God. By the way, I went to a catholic school for my first eight years and was taught evolution in science class. I guess if some kids are home schooled, they could have this kind of science hidden from them. But I believe they need to pass qualification testing in science to move on and graduate. I don't know of any schools that don't teach evolution.
I understand. At the same time though, if you literally see a clock's parts evolve into a clock one gear at a time through a process of trial and error-- and you had the finished product to refer to, and you had time capsules of the transitionary phases of past clock's progression into the complex machines that we use now (which we are still observing evolving to this day)--and the nuts and bolts and gears were following the same progression that you've been able to observe in already formed clocks--then would you still say that those gears and nuts and bolts wont eventually become a clock if they keep following the same pattern of formation based upon observed universal laws of physics??
(Bear with me if you please, I'm tired )
Lets say clocks represent a human body, kogs/gears/bolts represent the building blocks of the body, and the process of trial and error is evolution.
What's your opinion about the speciation proof I presented?
Lastly, I was in high school from 2001-2004. I remember a lot of hubub in the news about what to teach and not to teach in biology in Kansas schools. The only word I heard on evolution was that the courts mandated that that was the official version of things. To stay as far away out of controversy for fear of legal ramifications--the teacher got to the point that all we did in biology was cut open dead baby pigs and frogs and study mitosis and mitochondrial processes. Never heard a word on evolution or the fossil record. Genetics was condensed down to eye and hair color, genetic disease, and blood types. That's.. that's all I learned in biology.
I went on to Chemistry and physics from there. Admittedly I was private or homeschooled for all of middle school. But I have asked my friends from high school if they learned evolution in school before 9th grade and they said no. Anyone I know who hasn't been to college, have zero knowledge of evolution and are typically heavily biased against it.
If I were to create self aware beings knowing fully what they would do in their lifetimes, I sure wouldn't create a HELL for the majority of them to live in infinitely! That's not Love, that's sadistic. Therefore a truly loving god does not exist!
Quote:The sin is against an infinite being (God) unforgiven infinitely, therefore the punishment is infinite.
Dead wrong. The actions of a finite being measured against an infinite one are infinitesimal and therefore merit infinitesimal punishment.
Quote:Some people deserve hell.
I say again: No exceptions. Punishment should be equal to the crime, not in excess of it. As soon as the punishment is greater than the crime, the punisher is in the wrong.
(July 13, 2014 at 12:42 am)alexwenzel Wrote: "not one change of species into another is on record. We can not prove that a SINGLE specie has ever changed". That quote is by Charles Darwin himself. hock:
That statement still holds true today.
Yeah about Darwin now that we are a boarding the subject. There is one thing that has been bothering me, and I was wondering if anyone can give me some info about that.
(July 12, 2014 at 7:03 am)BrianSoddingBoru4 Wrote: 'Darwin was a racist, so evolution can't be true'
Is this true? I'm not talking about the evolution true stuff.
Because I've never heard that until know.
And if someone could answer the whole "Was Darwin a racist?" could you add some citations please.
July 13, 2014 at 1:46 am (This post was last modified: July 13, 2014 at 2:11 am by The Grand Nudger.)
He was a man of his time. Yep. He was also an abolitionist. People can be complicated. Try The Descent of Man.
Bear in mind, that at his time the word "racist" as we use it today we simply sound absurd. There were races of plants. Not people. He constantly made the case that with regards to people, we shouldn't really think in terms of race (species) - that we were all one species. Nevertheless, there is the assumption of western european (read:white) superiority just about everywhere in his comments with regards to what we would call races. He just didn't think it was a measure of biology.
(there's a tendency to "mythicize" important people and their positions on issues that resonate with us today, for better or worse - so while one side might say that darwinism led to eugenics and nazism - it didn't; the other would claim that evolutionary theory dispelled the myth of white supremecy - it didn't)
I am the Infantry. I am my country’s strength in war, her deterrent in peace. I am the heart of the fight… wherever, whenever. I carry America’s faith and honor against her enemies. I am the Queen of Battle. I am what my country expects me to be, the best trained Soldier in the world. In the race for victory, I am swift, determined, and courageous, armed with a fierce will to win. Never will I fail my country’s trust. Always I fight on…through the foe, to the objective, to triumph overall. If necessary, I will fight to my death. By my steadfast courage, I have won more than 200 years of freedom. I yield not to weakness, to hunger, to cowardice, to fatigue, to superior odds, For I am mentally tough, physically strong, and morally straight. I forsake not, my country, my mission, my comrades, my sacred duty. I am relentless. I am always there, now and forever. I AM THE INFANTRY! FOLLOW ME!
@Zidneya -These poor xtards don't want any help. https://plus.google.com/b/11256383813240...npAiTkrTTU
Funny how he keeps repeating "There's evidence for God all around you", but later in the conversation does not prevent any evidence. Rather, he defends himself with the claim that "the evolutionists are too deluded with the facts of evolution to see it". Oh, and this I learned something new today: evolution is indoctrination. Yes. Definitely not education.
(July 13, 2014 at 12:42 am)alexwenzel Wrote: "not one change of species into another is on record. We can not prove that a SINGLE specie has ever changed". That quote is by Charles Darwin himself. hock:
That statement still holds true today.
Yeah about Darwin now that we are a boarding the subject. There is one thing that has been bothering me, and I was wondering if anyone can give me some info about that.
(July 12, 2014 at 7:03 am)BrianSoddingBoru4 Wrote: 'Darwin was a racist, so evolution can't be true'
Is this true? I'm not talking about the evolution true stuff.
Because I've never heard that until know.
And if someone could answer the whole "Was Darwin a racist?" could you add some citations please.
No, of course Darwin wasn't a racist. But this is one of the non-arguments used by the slackwits to 'disprove' evolution. What they fail to grasp (or refuse to grasp) is that evolution stands on its own merits, regardless of Darwin's views on race. Odd, innit, how they never mention the blatant, flagrant racism of creationist Louis Agassiz?
Boru
‘I can’t be having with this.’ - Esmeralda Weatherwax
(July 13, 2014 at 6:17 am)BrianSoddingBoru4 Wrote: Odd, innit, how they never mention the blatant, flagrant racism of creationist Louis Agassiz?
Or the racism in the bible, which puts one race on a pedestal as the chosen race, and tells the believers that everyone else is fit for slavery, or murder... whatever. The best anyone else can hope for is to be left alone, assuming they don't get in Israel's way.
Not like the New Testament repeals that either, what with Jesus calling every other race dogs...
"YOU take the hard look in the mirror. You are everything that is wrong with this world. The only thing important to you, is you." - ronedee
Want to see more of my writing? Check out my (safe for work!) site, Unprotected Sects!
July 13, 2014 at 8:51 am (This post was last modified: July 13, 2014 at 8:51 am by Thumpalumpacus.)
(July 13, 2014 at 12:42 am)alexwenzel Wrote: "not one change of species into another is on record. We can not prove that a SINGLE specie has ever changed". That quote is by Charles Darwin himself. hock:
That statement still holds true today.
I have emphasized the portion of this statement which is factually incorrect.
For further reading:
Quote:5.2 Speciations in Plant Species not Involving Hybridization or Polyploidy
5.2.1 Stephanomeira malheurensis
Gottlieb (1973) documented the speciation of Stephanomeira malheurensis. He found a single small population (< 250 plants) among a much larger population (> 25,000 plants) of S. exigua in Harney Co., Oregon. Both species are diploid and have the same number of chromosomes (N = 8). S. exigua is an obligate outcrosser exhibiting sporophytic self-incompatibility. S. malheurensis exhibits no self-incompatibility and self-pollinates. Though the two species look very similar, Gottlieb was able to document morphological differences in five characters plus chromosomal differences. F1 hybrids between the species produces only 50% of the seeds and 24% of the pollen that conspecific crosses produced. F2 hybrids showed various developmental abnormalities.
5.2.2 Maize (Zea mays)
Pasterniani (1969) produced almost complete reproductive isolation between two varieties of maize. The varieties were distinguishable by seed color, white versus yellow. Other genetic markers allowed him to identify hybrids. The two varieties were planted in a common field. Any plant's nearest neighbors were always plants of the other strain. Selection was applied against hybridization by using only those ears of corn that showed a low degree of hybridization as the source of the next years seed. Only parental type kernels from these ears were planted. The strength of selection was increased each year. In the first year, only ears with less than 30% intercrossed seed were used. In the fifth year, only ears with less than 1% intercrossed seed were used. After five years the average percentage of intercrossed matings dropped from 35.8% to 4.9% in the white strain and from 46.7% to 3.4% in the yellow strain.
5.2.3 Speciation as a Result of Selection for Tolerance to a Toxin: Yellow Monkey Flower (Mimulus guttatus)
At reasonably low concentrations, copper is toxic to many plant species. Several plants have been seen to develop a tolerance to this metal (Macnair 1981). Macnair and Christie (1983) used this to examine the genetic basis of a postmating isolating mechanism in yellow monkey flower. When they crossed plants from the copper tolerant "Copperopolis" population with plants from the nontolerant "Cerig" population, they found that many of the hybrids were inviable. During early growth, just after the four leaf stage, the leaves of many of the hybrids turned yellow and became necrotic. Death followed this. This was seen only in hybrids between the two populations. Through mapping studies, the authors were able to show that the copper tolerance gene and the gene responsible for hybrid inviability were either the same gene or were very tightly linked. These results suggest that reproductive isolation may require changes in only a small number of genes.
5.3 The Fruit Fly Literature
5.3.1 Drosophila paulistorum
Dobzhansky and Pavlovsky (1971) reported a speciation event that occurred in a laboratory culture of Drosophila paulistorum sometime between 1958 and 1963. The culture was descended from a single inseminated female that was captured in the Llanos of Colombia. In 1958 this strain produced fertile hybrids when crossed with conspecifics of different strains from Orinocan. From 1963 onward crosses with Orinocan strains produced only sterile males. Initially no assortative mating or behavioral isolation was seen between the Llanos strain and the Orinocan strains. Later on Dobzhansky produced assortative mating (Dobzhansky 1972).
5.3.2 Disruptive Selection on Drosophila melanogaster
Thoday and Gibson (1962) established a population of Drosophila melanogaster from four gravid females. They applied selection on this population for flies with the highest and lowest numbers of sternoplural chaetae (hairs). In each generation, eight flies with high numbers of chaetae were allowed to interbreed and eight flies with low numbers of chaetae were allowed to interbreed. Periodically they performed mate choice experiments on the two lines. They found that they had produced a high degree of positive assortative mating between the two groups. In the decade or so following this, eighteen labs attempted unsuccessfully to reproduce these results. References are given in Thoday and Gibson 1970.
5.3.3 Selection on Courtship Behavior in Drosophila melanogaster
Crossley (1974) was able to produce changes in mating behavior in two mutant strains of D. melanogaster. Four treatments were used. In each treatment, 55 virgin males and 55 virgin females of both ebony body mutant flies and vestigial wing mutant flies (220 flies total) were put into a jar and allowed to mate for 20 hours. The females were collected and each was put into a separate vial. The phenotypes of the offspring were recorded. Wild type offspring were hybrids between the mutants. In two of the four treatments, mating was carried out in the light. In one of these treatments all hybrid offspring were destroyed. This was repeated for 40 generations. Mating was carried out in the dark in the other two treatments. Again, in one of these all hybrids were destroyed. This was repeated for 49 generations. Crossley ran mate choice tests and observed mating behavior. Positive assortative mating was found in the treatment which had mated in the light and had been subject to strong selection against hybridization. The basis of this was changes in the courtship behaviors of both sexes. Similar experiments, without observation of mating behavior, were performed by Knight, et al. (1956).
5.3.4 Sexual Isolation as a Byproduct of Adaptation to Environmental Conditions in Drosophila melanogaster
Kilias, et al. (1980) exposed D. melanogaster populations to different temperature and humidity regimes for several years. They performed mating tests to check for reproductive isolation. They found some sterility in crosses among populations raised under different conditions. They also showed some positive assortative mating. These things were not observed in populations which were separated but raised under the same conditions. They concluded that sexual isolation was produced as a byproduct of selection.
5.3.5 Sympatric Speciation in Drosophila melanogaster
In a series of papers (Rice 1985, Rice and Salt 1988 and Rice and Salt 1990) Rice and Salt presented experimental evidence for the possibility of sympatric speciation. They started from the premise that whenever organisms sort themselves into the environment first and then mate locally, individuals with the same habitat preferences will necessarily mate assortatively. They established a stock population of D. melanogaster with flies collected in an orchard near Davis, California. Pupae from the culture were placed into a habitat maze. Newly emerged flies had to negotiate the maze to find food. The maze simulated several environmental gradients simultaneously. The flies had to make three choices of which way to go. The first was between light and dark (phototaxis). The second was between up and down (geotaxis). The last was between the scent of acetaldehyde and the scent of ethanol (chemotaxis). This divided the flies among eight habitats. The flies were further divided by the time of day of emergence. In total the flies were divided among 24 spatio-temporal habitats.
They next cultured two strains of flies that had chosen opposite habitats. One strain emerged early, flew upward and was attracted to dark and acetaldehyde. The other emerged late, flew downward and was attracted to light and ethanol. Pupae from these two strains were placed together in the maze. They were allowed to mate at the food site and were collected. Eye color differences between the strains allowed Rice and Salt to distinguish between the two strains. A selective penalty was imposed on flies that switched habitats. Females that switched habitats were destroyed. None of their gametes passed into the next generation. Males that switched habitats received no penalty. After 25 generations of this mating tests showed reproductive isolation between the two strains. Habitat specialization was also produced.
They next repeated the experiment without the penalty against habitat switching. The result was the same -- reproductive isolation was produced. They argued that a switching penalty is not necessary to produce reproductive isolation. Their results, they stated, show the possibility of sympatric speciation.
5.3.6 Isolation Produced as an Incidental Effect of Selection on several Drosophila species
In a series of experiments, del Solar (1966) derived positively and negatively geotactic and phototactic strains of D. pseudoobscura from the same population by running the flies through mazes. Flies from different strains were then introduced into mating chambers (10 males and 10 females from each strain). Matings were recorded. Statistically significant positive assortative mating was found.
In a separate series of experiments Dodd (1989) raised eight populations derived from a single population of D. Pseudoobscura on stressful media. Four populations were raised on a starch based medium, the other four were raised on a maltose based medium. The fly populations in both treatments took several months to get established, implying that they were under strong selection. Dodd found some evidence of genetic divergence between flies in the two treatments. He performed mate choice tests among experimental populations. He found statistically significant assortative mating between populations raised on different media, but no assortative mating among populations raised within the same medium regime. He argued that since there was no direct selection for reproductive isolation, the behavioral isolation results from a pleiotropic by-product to adaptation to the two media. Schluter and Nagel (1995) have argued that these results provide experimental support for the hypothesis of parallel speciation.
Less dramatic results were obtained by growing D. willistoni on media of different pH levels (de Oliveira and Cordeiro 1980). Mate choice tests after 26, 32, 52 and 69 generations of growth showed statistically significant assortative mating between some populations grown in different pH treatments. This ethological isolation did not always persist over time. They also found that some crosses made after 106 and 122 generations showed significant hybrid inferiority, but only when grown in acid medium.
5.3.7 Selection for Reinforcement in Drosophila melanogaster
Some proposed models of speciation rely on a process called reinforcement to complete the speciation process. Reinforcement occurs when to partially isolated allopatric populations come into contact. Lower relative fitness of hybrids between the two populations results in increased selection for isolating mechanisms. I should note that a recent review (Rice and Hostert 1993) argues that there is little experimental evidence to support reinforcement models. Two experiments in which the authors argue that their results provide support are discussed below.
Ehrman (1971) established strains of wild-type and mutant (black body) D. melanogaster. These flies were derived from compound autosome strains such that heterotypic matings would produce no progeny. The two strains were reared together in common fly cages. After two years, the isolation index generated from mate choice experiments had increased from 0.04 to 0.43, indicating the appearance of considerable assortative mating. After four years this index had risen to 0.64 (Ehrman 1973).
Along the same lines, Koopman (1950) was able to increase the degree of reproductive isolation between two partially isolated species, D. pseudoobscura and D. persimilis.
5.3.8 Tests of the Founder-flush Speciation Hypothesis Using Drosophila
The founder-flush (a.k.a. flush-crash) hypothesis posits that genetic drift and founder effects play a major role in speciation (Powell 1978). During a founder-flush cycle a new habitat is colonized by a small number of individuals (e.g. one inseminated female). The population rapidly expands (the flush phase). This is followed by the population crashing. During this crash period the population experiences strong genetic drift. The population undergoes another rapid expansion followed by another crash. This cycle repeats several times. Reproductive isolation is produced as a byproduct of genetic drift.
Dodd and Powell (1985) tested this hypothesis using D. pseudoobscura. A large, heterogeneous population was allowed to grow rapidly in a very large population cage. Twelve experimental populations were derived from this population from single pair matings. These populations were allowed to flush. Fourteen months later, mating tests were performed among the twelve populations. No postmating isolation was seen. One cross showed strong behavioral isolation. The populations underwent three more flush-crash cycles. Forty-four months after the start of the experiment (and fifteen months after the last flush) the populations were again tested. Once again, no postmating isolation was seen. Three populations showed behavioral isolation in the form of positive assortative mating. Later tests between 1980 and 1984 showed that the isolation persisted, though it was weaker in some cases.
Galina, et al. (1993) performed similar experiments with D. pseudoobscura. Mating tests between populations that underwent flush-crash cycles and their ancestral populations showed 8 cases of positive assortative mating out of 118 crosses. They also showed 5 cases of negative assortative mating (i.e. the flies preferred to mate with flies of the other strain). Tests among the founder-flush populations showed 36 cases of positive assortative mating out of 370 crosses. These tests also found 4 cases of negative assortative mating. Most of these mating preferences did not persist over time. Galina, et al. concluded that the founder-flush protocol yields reproductive isolation only as a rare and erratic event.
Ahearn (1980) applied the founder-flush protocol to D. silvestris. Flies from a line of this species underwent several flush-crash cycles. They were tested in mate choice experiments against flies from a continuously large population. Female flies from both strains preferred to mate with males from the large population. Females from the large population would not mate with males from the founder flush population. An asymmetric reproductive isolation was produced.
In a three year experiment, Ringo, et al. (1985) compared the effects of a founder-flush protocol to the effects of selection on various traits. A large population of D. simulans was created from flies from 69 wild caught stocks from several locations. Founder-flush lines and selection lines were derived from this population. The founder-flush lines went through six flush-crash cycles. The selection lines experienced equal intensities of selection for various traits. Mating test were performed between strains within a treatment and between treatment strains and the source population. Crosses were also checked for postmating isolation. In the selection lines, 10 out of 216 crosses showed positive assortative mating (2 crosses showed negative assortative mating). They also found that 25 out of 216 crosses showed postmating isolation. Of these, 9 cases involved crosses with the source population. In the founder-flush lines 12 out of 216 crosses showed positive assortative mating (3 crosses showed negative assortative mating). Postmating isolation was found in 15 out of 216 crosses, 11 involving the source population. They concluded that only weak isolation was found and that there was little difference between the effects of natural selection and the effects of genetic drift.
A final test of the founder-flush hypothesis will be described with the housefly cases below.
5.4 Housefly Speciation Experiments
5.4.1 A Test of the Founder-flush Hypothesis Using Houseflies
Meffert and Bryant (1991) used houseflies to test whether bottlenecks in populations can cause permanent alterations in courtship behavior that lead to premating isolation. They collected over 100 flies of each sex from a landfill near Alvin, Texas. These were used to initiate an ancestral population. From this ancestral population they established six lines. Two of these lines were started with one pair of flies, two lines were started with four pairs of flies and two lines were started with sixteen pairs of flies. These populations were flushed to about 2,000 flies each. They then went through five bottlenecks followed by flushes. This took 35 generations. Mate choice tests were performed. One case of positive assortative mating was found. One case of negative assortative mating was also found.
5.4.2 Selection for Geotaxis with and without Gene Flow
Soans, et al. (1974) used houseflies to test Pimentel's model of speciation. This model posits that speciation requires two steps. The first is the formation of races in subpopulations. This is followed by the establishment of reproductive isolation. Houseflies were subjected to intense divergent selection on the basis of positive and negative geotaxis. In some treatments no gene flow was allowed, while in others there was 30% gene flow. Selection was imposed by placing 1000 flies into the center of a 108 cm vertical tube. The first 50 flies that reached the top and the first 50 flies that reached the bottom were used to found positively and negatively geotactic populations. Four populations were established:
Population A + geotaxis, no gene flow
Population B - geotaxis, no gene flow
Population C + geotaxis, 30% gene flow
Population D - geotaxis, 30% gene flow
Selection was repeated within these populations each generations. After 38 generations the time to collect 50 flies had dropped from 6 hours to 2 hours in Pop A, from 4 hours to 4 minutes in Pop B, from 6 hours to 2 hours in Pop C and from 4 hours to 45 minutes in Pop D. Mate choice tests were performed. Positive assortative mating was found in all crosses. They concluded that reproductive isolation occurred under both allopatric and sympatric conditions when very strong selection was present.
Hurd and Eisenberg (1975) performed a similar experiment on houseflies using 50% gene flow and got the same results.
5.5 Speciation Through Host Race Differentiation
Recently there has been a lot of interest in whether the differentiation of an herbivorous or parasitic species into races living on different hosts can lead to sympatric speciation. It has been argued that in animals that mate on (or in) their preferred hosts, positive assortative mating is an inevitable byproduct of habitat selection (Rice 1985; Barton, et al. 1988). This would suggest that differentiated host races may represent incipient species.
5.5.1 Apple Maggot Fly (Rhagoletis pomonella)
Rhagoletis pomonella is a fly that is native to North America. Its normal host is the hawthorn tree. Sometime during the nineteenth century it began to infest apple trees. Since then it has begun to infest cherries, roses, pears and possibly other members of the rosaceae. Quite a bit of work has been done on the differences between flies infesting hawthorn and flies infesting apple. There appear to be differences in host preferences among populations. Offspring of females collected from on of these two hosts are more likely to select that host for oviposition (Prokopy et al. 1988). Genetic differences between flies on these two hosts have been found at 6 out of 13 allozyme loci (Feder et al. 1988, see also McPheron et al. 1988). Laboratory studies have shown an asynchrony in emergence time of adults between these two host races (Smith 1988). Flies from apple trees take about 40 days to mature, whereas flies from hawthorn trees take 54-60 days to mature. This makes sense when we consider that hawthorn fruit tends to mature later in the season that apples. Hybridization studies show that host preferences are inherited, but give no evidence of barriers to mating. This is a very exciting case. It may represent the early stages of a sympatric speciation event (considering the dispersal of R. pomonella to other plants it may even represent the beginning of an adaptive radiation). It is important to note that some of the leading researchers on this question are urging caution in interpreting it. Feder and Bush (1989) stated:
"Hawthorn and apple "host races" of R. pomonella may therefore represent incipient species. However, it remains to be seen whether host-associated traits can evolve into effective enough barriers to gene flow to result eventually in the complete reproductive isolation of R. pomonella populations."
5.5.2 Gall Former Fly (Eurosta solidaginis)
Eurosta solidaginis is a gall forming fly that is associated with goldenrod plants. It has two hosts: over most of its range it lays its eggs in Solidago altissima, but in some areas it uses S. gigantea as its host. Recent electrophoretic work has shown that the genetic distances among flies from different sympatric hosts species are greater than the distances among flies on the same host in different geographic areas (Waring et al. 1990). This same study also found reduced variability in flies on S. gigantea. This suggests that some E. solidaginis have recently shifted hosts to this species. A recent study has compared reproductive behavior of the flies associated with the two hosts (Craig et al. 1993). They found that flies associated with S. gigantea emerge earlier in the season than flies associated with S. altissima. In host choice experiments, each fly strain ovipunctured its own host much more frequently than the other host. Craig et al. (1993) also performed several mating experiments. When no host was present and females mated with males from either strain, if males from only one strain were present. When males of both strains were present, statistically significant positive assortative mating was seen. In the presence of a host, assortative mating was also seen. When both hosts and flies from both populations were present, females waited on the buds of the host that they are normally associated with. The males fly to the host to mate. Like the Rhagoletis case above, this may represent the beginning of a sympatric speciation.
5.6 Flour Beetles (Tribolium castaneum)
Halliburton and Gall (1981) established a population of flour beetles collected in Davis, California. In each generation they selected the 8 lightest and the 8 heaviest pupae of each sex. When these 32 beetles had emerged, they were placed together and allowed to mate for 24 hours. Eggs were collected for 48 hours. The pupae that developed from these eggs were weighed at 19 days. This was repeated for 15 generations. The results of mate choice tests between heavy and light beetles was compared to tests among control lines derived from randomly chosen pupae. Positive assortative mating on the basis of size was found in 2 out of 4 experimental lines.
5.7 Speciation in a Lab Rat Worm, Nereis acuminata
In 1964 five or six individuals of the polychaete worm, Nereis acuminata, were collected in Long Beach Harbor, California. These were allowed to grow into a population of thousands of individuals. Four pairs from this population were transferred to the Woods Hole Oceanographic Institute. For over 20 years these worms were used as test organisms in environmental toxicology. From 1986 to 1991 the Long Beach area was searched for populations of the worm. Two populations, P1 and P2, were found. Weinberg, et al. (1992) performed tests on these two populations and the Woods Hole population (WH) for both postmating and premating isolation. To test for postmating isolation, they looked at whether broods from crosses were successfully reared. The results below give the percentage of successful rearings for each group of crosses.
WH × WH - 75%
P1 × P1 - 95%
P2 × P2 - 80%
P1 × P2 - 77%
WH × P1 - 0%
WH × P2 - 0%
They also found statistically significant premating isolation between the WH population and the field populations. Finally, the Woods Hole population showed slightly different karyotypes from the field populations.
5.8 Speciation Through Cytoplasmic Incompatability Resulting from the Presence of a Parasite or Symbiont
In some species the presence of intracellular bacterial parasites (or symbionts) is associated with postmating isolation. This results from a cytoplasmic incompatability between gametes from strains that have the parasite (or symbiont) and stains that don't. An example of this is seen in the mosquito Culex pipiens (Yen and Barr 1971). Compared to within strain matings, matings between strains from different geographic regions may may have any of three results: These matings may produce a normal number of offspring, they may produce a reduced number of offspring or they may produce no offspring. Reciprocal crosses may give the same or different results. In an incompatible cross, the egg and sperm nuclei fail to unite during fertilization. The egg dies during embryogenesis. In some of these strains, Yen and Barr (1971) found substantial numbers of Rickettsia-like microbes in adults, eggs and embryos. Compatibility of mosquito strains seems to be correlated with the strain of the microbe present. Mosquitoes that carry different strains of the microbe exhibit cytoplasmic incompatibility; those that carry the same strain of microbe are interfertile.
Similar phenomena have been seen in a number of other insects. Microoganisms are seen in the eggs of both Nasonia vitripennis and N. giraulti. These two species do not normally hybridize. Following treatment with antibiotics, hybrids occur between them (Breeuwer and Werren 1990). In this case, the symbiont is associated with improper condensation of host chromosomes.
For more examples and a critical review of this topic, see Thompson 1987.
July 13, 2014 at 10:33 am (This post was last modified: July 13, 2014 at 10:35 am by The Grand Nudger.)
(July 13, 2014 at 6:17 am)BrianSoddingBoru4 Wrote: No, of course Darwin wasn't a racist.
Boru
He was far less racist than his peers, sure. That doesn't mean he wasn't a racist. My grandfather is less racist than a klansman or a neo-nazi (he thinks both groups of people are ignorant bastards). He's still a racist. In fact, he's a racist in the pretty much the same way that Darwin was.
I am the Infantry. I am my country’s strength in war, her deterrent in peace. I am the heart of the fight… wherever, whenever. I carry America’s faith and honor against her enemies. I am the Queen of Battle. I am what my country expects me to be, the best trained Soldier in the world. In the race for victory, I am swift, determined, and courageous, armed with a fierce will to win. Never will I fail my country’s trust. Always I fight on…through the foe, to the objective, to triumph overall. If necessary, I will fight to my death. By my steadfast courage, I have won more than 200 years of freedom. I yield not to weakness, to hunger, to cowardice, to fatigue, to superior odds, For I am mentally tough, physically strong, and morally straight. I forsake not, my country, my mission, my comrades, my sacred duty. I am relentless. I am always there, now and forever. I AM THE INFANTRY! FOLLOW ME!